Ferns of Thailand, Laos and Cambodia

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Family

Thelypteridaceae

Nomenclature

Cyclosorus papilio (C.Hope) Ching, Bull. Fan Mem. Inst. Biol. 8: 214. 1938; Tardieu & C.Chr., Fl. Indo-Chine 7(2) 382. 1941; Holttum, Rev. Fl. Malaya, ed. 2, 2: 633. 1968. – Nephrodium papilio C.Hope, J. Bomb. Nat. Hist. Soc. 12: 625, t. 12. 1899. – Thelypteris papilio (C.Hope) K.Iwats., Mem. Coll. Sci. Univ. Kyoto B. 31: 175. 1965; Tagawa & K.Iwats., SouthE. Asian Stud. 3(3): 79. 1965; Tagawa & K.Iwats., Fl. Thailand 3: 428. 1988. – Christella papilio (C.Hope) Holttum in B.K.Nayar & S.Kaur, Companion Handb. Ferns Brit. India: 208. 1974; Holttum, Kew Bull. 31: 321. 1976; Holttum, Fl. Males., Ser. II, Pterid. 1: 556, f. 20d–e. 1982 [‘1981’]; Boonkerd & Pollawatn, Pterid. Thailand: 221. 2000. – Lectotype (Holttum, Kew Bulletin, 31(2); 322. 1976): Levigne s.n. 1880, N.E. India, below Darjeeling 1200 m (K).

Description

Description adapted from Holttum, Fl. Males., Ser. II, Pterid. 1: 556, f. 20d–e. 1982 [‘1981’].
Rhizome massive, erect; stipe 10–30 cm long, minutely hairy, basal scales thin, c. 7 x 1 mm. Laminae to 150 cm or more long, to 36 cm wide; up to 30 pairs of pinnae, the lowermost c. 10 pairs gradually decrescent, broadly triangular, strongly auricled on acroscopic base, broadly truncate on basiscopic, margins above base shallowly lobed, apex acuminate; lowest pinnae 1–3 cm long. Largest pinnae to 17 x 2.5 cm; base subtruncate; apex acuminate with short cauda; edges lobed 1/4–1/3 towards costa (most deeply in fertile pinnae), lobes slightly falcate and rounded at their tips; costules 4–4.5 mm apart; veins 7–9 pairs, 1.5 pairs anastomosing, 1–2 pairs ending beside the sinus-membrane; lower surface of all parts bearing very short erect hairs (less than 0.1 mm long on rachis and bases of costae, a little longer on distal parts of pinnae), some thick orange glandular hairs sometimes present on veins in pinna-lobes; upper surface of costae covered with hairs 0.3–0.5 mm long, scattered shorter hairs on costules, minute erect hairs on surface between veins. Sori medial; indusia rather large, thin, with short hairs as lamina .

Distribution in Thailand

NORTHERN: Chiang Mai, Tak, Phitsanulok; SOUTH-WESTERN: Kanchanaburi, Prachuap Khiri Khan.

Wider Distribution

Sri Lanka, Himalaya, Taiwan, and Peninsular Malaysia.

Ecology

On moist ground by streamlets at edge of evergreen forests at low to high altitudes.

Proposed IUCN Conservation Assessment

Least Concern (LC). This species is widespread and not under any known threat.

Voucher specimens - Thailand

Smith 2725, Tak, Raheng (K); Shimizu et al. T-11616, Phitsanulok, Phu Miang (K).

	Ferns of Thailand, Laos and Cambodia
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Cyclosorus papilio (C.Hope) Ching Warning: count(): Parameter must be an array or an object that implements Countable in C:\Data\serverroots\htdocs\ThaiFerns\factsheets\functions.php on line 17 Family Thelypteridaceae Nomenclature Cyclosorus papilio (C.Hope) Ching, Bull. Fan Mem. Inst. Biol. 8: 214. 1938; Tardieu & C.Chr., Fl. Indo-Chine 7(2) 382. 1941; Holttum, Rev. Fl. Malaya, ed. 2, 2: 633. 1968. – Nephrodium papilio C.Hope, J. Bomb. Nat. Hist. Soc. 12: 625, t. 12. 1899. – Thelypteris papilio (C.Hope) K.Iwats., Mem. Coll. Sci. Univ. Kyoto B. 31: 175. 1965; Tagawa & K.Iwats., SouthE. Asian Stud. 3(3): 79. 1965; Tagawa & K.Iwats., Fl. Thailand 3: 428. 1988. – Christella papilio (C.Hope) Holttum in B.K.Nayar & S.Kaur, Companion Handb. Ferns Brit. India: 208. 1974; Holttum, Kew Bull. 31: 321. 1976; Holttum, Fl. Males., Ser. II, Pterid. 1: 556, f. 20d–e. 1982 [‘1981’]; Boonkerd & Pollawatn, Pterid. Thailand: 221. 2000. – Lectotype (Holttum, Kew Bulletin, 31(2); 322. 1976): Levigne s.n. 1880, N.E. India, below Darjeeling 1200 m (K). Description Description adapted from Holttum, Fl. Males., Ser. II, Pterid. 1: 556, f. 20d–e. 1982 [‘1981’]. Rhizome massive, erect; stipe 10–30 cm long, minutely hairy, basal scales thin, c. 7 x 1 mm. Laminae to 150 cm or more long, to 36 cm wide; up to 30 pairs of pinnae, the lowermost c. 10 pairs gradually decrescent, broadly triangular, strongly auricled on acroscopic base, broadly truncate on basiscopic, margins above base shallowly lobed, apex acuminate; lowest pinnae 1–3 cm long. Largest pinnae to 17 x 2.5 cm; base subtruncate; apex acuminate with short cauda; edges lobed 1/4–1/3 towards costa (most deeply in fertile pinnae), lobes slightly falcate and rounded at their tips; costules 4–4.5 mm apart; veins 7–9 pairs, 1.5 pairs anastomosing, 1–2 pairs ending beside the sinus-membrane; lower surface of all parts bearing very short erect hairs (less than 0.1 mm long on rachis and bases of costae, a little longer on distal parts of pinnae), some thick orange glandular hairs sometimes present on veins in pinna-lobes; upper surface of costae covered with hairs 0.3–0.5 mm long, scattered shorter hairs on costules, minute erect hairs on surface between veins. Sori medial; indusia rather large, thin, with short hairs as lamina . Distribution in Thailand NORTHERN: Chiang Mai, Tak, Phitsanulok; SOUTH-WESTERN: Kanchanaburi, Prachuap Khiri Khan. Wider Distribution Sri Lanka, Himalaya, Taiwan, and Peninsular Malaysia. Ecology On moist ground by streamlets at edge of evergreen forests at low to high altitudes. Proposed IUCN Conservation Assessment Least Concern (LC). This species is widespread and not under any known threat. Voucher specimens - Thailand Smith 2725, Tak, Raheng (K); Shimizu et al. T-11616, Phitsanulok, Phu Miang (K). Warning: count(): Parameter must be an array or an object that implements Countable in C:\Data\serverroots\htdocs\ThaiFerns\factsheets\functions.php on line 17
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Cyclosorus papilio (C.Hope) Ching