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Davallia heterophylla Sm.FamilyDavalliaceae NomenclatureDavallia heterophylla Sm., Mém. Acad. Roy. Sci. (Turin) 5: 415. 1793; Tagawa & K.Iwats., Fl. Thailand 3: 616. 1989; Nooteboom, Fl. Males., Ser. II, Ferns and Fern Allies 3: 254. 1998. – Humata heterophylla (Sm.) Desv., Mém. Soc. Linn. Paris 6(3): 323. 1827; Bedd, Handb. Ferns Brit. India: 46. 1883; Bonap., Notes Ptérid. 14: 53. 1923; E.Smith, J. Siam Soc. Nat. Hist. Suppl. 8: 2. 1929; Tardieu & C.Chr., Fl. Indo-Chine 7(2): 109. 1939; Holttum, Rev. Fl. Malaya ed. 1, 2: 366, f. 211. 1955 [‘1954’]; Tagawa & K.Iwats., Fl. Thailand 3: 168. 1985; Boonkerd & Pollawatn, Pterid. Thailand: 171, 235. 2000. – Type: Charles Miller 1778 (BM). DescriptionRhizome long creeping, about 1.5 mm diam., bearing dimorphic fronds remotely, glaucous, densely scaly; scales gradually narrowing from oblong base towards long tails, up to 6 by 0.7 mm, tails usually patent, brown to darker, minutely toothed. Sterile frond: stipe 0.5–2.5 cm long, narrowly winged; lamina simple, oblong, acuminate at apex, round to cuneate at base, entire, 6–9 by 2–2.5 cm; coriaceous, naked; veins twice or thrice forked, all free. Fertile frond: stipe 1–2.5 cm long, winged; lamina linear-lanceolate, gradually narrowing towards both ends, up to 9 by 1 cm, lobed to half way; lobes patent, round, wavy. Sori terminal on veinlets, 3–5 for each lobe, a little inside the edges; indusia up to 1 mm long, 2 mm broad, brown, entire . Distribution in ThailandPENINSULAR: Surat Thani, Phangnga, Krabi, Nakhon Si Thammarat, Songkhla, Narathiwat. Wider DistributionThroughout Malesia to the Pacific, north to S Vietnam (“Cochinchina”). EcologyOn branches of trees in dense forests. Proposed IUCN Conservation AssessmentLeast Concern (LC). This species is widespread and not under any known threat. Voucher specimens - ThailandMiddleton et al. 5224, Songkhla, Ton Nga Chang Wildlife Sanctuary (E). Habit Habit Habit Habit Rhizome Sterile and fertile fronds Fertile and sterile fronds Variation from sterile (margin entire) to fertile fronds (margin lobed) and intermediate stages Sterile fronds Upper surface of fertile frond Lower surface of fertile frond Sori |
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