Family

Polypodiaceae

Nomenclature

Goniophlebium percussum (Cav.) Wagner & Grether, Occas. Pap. Bernice Pauahi Bishop Mus. 19: 88. 1948; Copeland, Fern Fl. Philipp. 3: 460. 1960; Rödl-Linder, Philipp. J. Sci. 116: 155. 1987; Rödl-Linder, Blumea 34: 381. 1990; Hovenkamp & Rödl-Linder, Fl. Males., Ser. II, Ferns and Fern Allies 3: 58. 1998. – Cyathea percussa Cav., Descr. Pl.: 548. 1802. – Schellolepis percussa (Cav.) Pic.Serm., Webbia 28: 470. 1973. – Polypodium cyathoides Sw., Syn. Fil.: 37. 1806, nom. nov., non Polypodium percussum Cav., C.Chr., Dansk Bot. Ark. 9: 29. 1937. – Goniophlebium cyathoides (Sw.) Hosok., Trans. Nat. Hist. Soc. Form . 32: 286. 1942. – Polypodium cyathoides forma typicum C.Chr., Ark. f. Bot. 9: 39. 1910, nom. illeg. – Type: Marianas, Née (MA, S-PA [fragment]).

Marginaria verrucosa Hook., Gen. Fil.: t. 14. 1838. – Goniophlebium verrucosum (Hook.) J.Sm. in Hook., Gen. Fil.: ad t. 51. 1840; Bedd., Handb. Ferns Brit. India: 324, f. 175. 1883; Boonkerd & Pollawatn, Pterid. Thailand: 272. 2000. – Polypodium verrucosum (Hook.) Wall. ex Hook., Gard. Ferns: t. 41. 1862; Holttum, Rev. Fl. Malaya ed. 1, 2: 206, f. 106. 1955 [‘1954’]; Tagawa & K.Iwats., Acta Phytotax. Geobot. 23: 53. 1968; Tagawa & K.Iwats., Fl. Thailand 3: 574. 1989. – Schellolepis verrucosa (Hook.) J.Sm., Ferns Brit. For.: 83. 1866.

Description

Description adapted from Hovenkamp & Rödl-Linder, Fl. Males., Ser. II, Ferns and Fern Allies 3: 58. 1998.
Rhizome long creeping, terete, (3–)5–11 mm thick, brown; phyllopodia 1.5–2.5 cm distant. Anatomy: vascular strands 14–19, with dark bundle sheaths, sclerenchyma strands many (70–100). Rhizome scales inserted evenly, deciduous, densely set, appressed, pseudopeltate, monomorphic, deltoid, 2.5–4 by 0.7–1.3 mm, index 2.8–3.6, basal auricles rounded, margin at the base sparsely set with glands, dentate, teeth all equally long or slightly shorter at base, apex acuminate to filiform, cell walls equally clathrate throughout, superficial hairs present, spread over the entire acumen. Fronds pinnate, stipe 3.5–5 mm thick, 0.2–0.6 x as long as the lamina, glabrous or minutely scaly; lamina 38–99(–200) by 18–38(–45) cm, index 2.1–4.4, widest at the base; pinnae articulate, shortly petiolate, 12.5–20.5 by 2–3 cm, index 6.2–6.8, base shortly angustate, margin entire to crenate, apex acuminate; basal pair 2–6 cm distant from next pair and equally long, not deflexed, apical segment conform, slightly longer than longest pinnae/segment. Venation: areoles 3– or 4-serial, marginal veinlets up to 1/3 as long as the areole, simple or forked, frequently anastomosing. Indument: 2-celled glandular hairs present (so small and sparse that lamina may appear glabrous), acicular hairs absent to dense all over the lamina, 2–5 cells long, scales deciduous. Sori uniserial, costal, deeply sunken, 0.8–1 mm diam., receptacular scaly paraphyses persistent, clathrate, 230–250 by 180–200 micrometres, 3 or more cells wide. Sporangia glabrous. Spores yellow, cristate , with short polar excrescences.

Distribution in Thailand

PENINSULAR: Nakhon Si Thammarat, Surat Thani, Yala, Narathiwat.

Wider Distribution

Throughout Malesia to Australia (Queensland).

Ecology

Epiphytic on tree trunks in low altitudes.

Proposed IUCN Conservation Assessment

Least Concern (LC). This species is widespread and not under any known threat.

Voucher specimens - Thailand

Suksathan 2953, Narathiwat (QBG).