Ferns of Thailand, Laos and Cambodia

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Family

Polypodiaceae

Nomenclature

Pyrrosia costata (C.Presl ex Bedd.) Tagawa & K.Iwats., Acta Phytotax. Geobot. 22: 100. 1967; Tagawa & K.Iwats., SouthE. Asian Stud. 5: 48. 1967; Hovenkamp, Leiden Bot. Ser. 9: 171, f. 17. 1986; Tagawa & K.Iwats., Fl. Thailand 3: 505. 1989; Boonkerd & Pollawatn, Pterid. Thailand: 285. 2000. – Niphobolus costatus C.Presl ex Bedd., Ferns Brit. India: t. 120. 1868, based on Polypodium costatum Wall. ex Mett., Abhandl. Senchkenb. Naturf. Ges. 2: 131. 1857, non Kunze 1834.

Niphobolus beddomeanus Giesenh., Nipholobus: 101. 1901. – Pyrrosia beddomeana (Giesenh.) Ching, Bull. Chin. Bot. Soc. 1: 68. 1935.

Description

Rhizome short creeping, bearing closely spaced fronds, about 3 mm diam., densely scaly at least at apex; scales linear-subulate, up to 1 mm broad at base, long-tailed, up to 1 cm long, more or less bi-coloured with dark brown to nearly black basal portion and brown to dark brown marginal portion and tails. Fronds simple. Stipes usually indistinct, the base of laminae long decurrent and winged nearly to the base, stramineous or pale brown, scaly at base, densely hairy throughout. Laminae broadly spathulate or oblanceolate, broadest at about 1/3 way from apex, caudately acuminate at apex, very gradually narrowing towards attenuate base and decurrent downwards to wings of stipes, the end not distinct, up to 40 cm long including indistinct stipes of 5–10 cm long, 2.5–5.5 cm broad, the sterile and the fertile fronds not different; midrib and main veins distinct, raised beneath, the other veins invisible, copiously anastomosing; thick but not stiff, upper surface deep green, stellate-hairy or glabrescent, with scattered hydathodes, lower surface densely covered with a mat of stellate hairs. Sori round , small but distinct, about 0.8 mm diam., usually covering the whole undersurface of fronds except for the lower portion.

Distribution in Thailand

NORTHERN: Chiang Mai, Chiang Rai; NORTH-EASTERN: Loei.

Wider Distribution

Himalayas, SW China (Yunnan, Tibet), Upper Burma and Vietnam.

Ecology

On rocks usually in light shade in evergreen forests at 800–1500 m alt., rather rare.

Proposed IUCN Conservation Assessment

Least Concern (LC). This species is widespread and not under any known threat.

	Ferns of Thailand, Laos and Cambodia
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Pyrrosia costata (C.Presl ex Bedd.) Tagawa & K.Iwats. Warning: count(): Parameter must be an array or an object that implements Countable in C:\Data\serverroots\htdocs\ThaiFerns\factsheets\functions.php on line 17 Family Polypodiaceae Nomenclature Pyrrosia costata (C.Presl ex Bedd.) Tagawa & K.Iwats., Acta Phytotax. Geobot. 22: 100. 1967; Tagawa & K.Iwats., SouthE. Asian Stud. 5: 48. 1967; Hovenkamp, Leiden Bot. Ser. 9: 171, f. 17. 1986; Tagawa & K.Iwats., Fl. Thailand 3: 505. 1989; Boonkerd & Pollawatn, Pterid. Thailand: 285. 2000. – Niphobolus costatus C.Presl ex Bedd., Ferns Brit. India: t. 120. 1868, based on Polypodium costatum Wall. ex Mett., Abhandl. Senchkenb. Naturf. Ges. 2: 131. 1857, non Kunze 1834. Niphobolus beddomeanus Giesenh., Nipholobus: 101. 1901. – Pyrrosia beddomeana (Giesenh.) Ching, Bull. Chin. Bot. Soc. 1: 68. 1935. Description Rhizome short creeping, bearing closely spaced fronds, about 3 mm diam., densely scaly at least at apex; scales linear-subulate, up to 1 mm broad at base, long-tailed, up to 1 cm long, more or less bi-coloured with dark brown to nearly black basal portion and brown to dark brown marginal portion and tails. Fronds simple. Stipes usually indistinct, the base of laminae long decurrent and winged nearly to the base, stramineous or pale brown, scaly at base, densely hairy throughout. Laminae broadly spathulate or oblanceolate, broadest at about 1/3 way from apex, caudately acuminate at apex, very gradually narrowing towards attenuate base and decurrent downwards to wings of stipes, the end not distinct, up to 40 cm long including indistinct stipes of 5–10 cm long, 2.5–5.5 cm broad, the sterile and the fertile fronds not different; midrib and main veins distinct, raised beneath, the other veins invisible, copiously anastomosing; thick but not stiff, upper surface deep green, stellate-hairy or glabrescent, with scattered hydathodes, lower surface densely covered with a mat of stellate hairs. Sori round , small but distinct, about 0.8 mm diam., usually covering the whole undersurface of fronds except for the lower portion. Distribution in Thailand NORTHERN: Chiang Mai, Chiang Rai; NORTH-EASTERN: Loei. Wider Distribution Himalayas, SW China (Yunnan, Tibet), Upper Burma and Vietnam. Ecology On rocks usually in light shade in evergreen forests at 800–1500 m alt., rather rare. Proposed IUCN Conservation Assessment Least Concern (LC). This species is widespread and not under any known threat. Warning: count(): Parameter must be an array or an object that implements Countable in C:\Data\serverroots\htdocs\ThaiFerns\factsheets\functions.php on line 17
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