Family

Tectaria impressa (Fée) Holttum

Nomenclature

Tectaria impressa (Fée) Holttum, Kew Bull. 43: 483. 1988; Tagawa & K.Iwats., Fl. Thailand 3: 621. 1988; Holttum, Fl. Males., Ser. II, Pterid. 2: 51. 1991; Boonkerd & Pollawatn, Pterid. Thailand: 162, 163, 218. 2000; Newman et al., Checkl. Vasc. Pl. Lao PDR: 32. 2007. – Phlebigonium impressum Fée, Gen. Fil.: 314. 1852. – Type:  Griffith s.n., Ind. Or. (RB).

Aspidium variolosum Wall. ex Hook., Sp. Fil. 4: 51. 1862; Bedd., Handb. Ferns Brit. India: 216, f. 111. 1883; Christ, Bot. Tidsskr. 24: 108. 1901; C.Chr., Bot. Tidsskr. 32: 419. 1916. – Nephrodium variolosum (Wall. ex Hook.) Hook. & Baker, Syn. Fil.: 298. 1867; Ostenfeld, Bull. Herb. Boiss. II. 5: 721. 1905. – Tectaria variolosa (Wall. ex Hook.) C.Chr., Contr. U.S. Natl. Herb. 26: 289. 1931; Ching, Sinensia 2: 21, pl. 3. 1931; Tardieu & C.Chr., Fl. Indo-Chine 7(2): 412. 1941; Holttum, Rev. Fl. Malaya ed. 1, 2: 506, f. 298. 1955 [‘1954’]; Holttum, Dansk Bot. Ark. 20: 31. 1961; Holttum, Dansk Bot. Ark 23: 242. 1965; Tagawa & K.Iwats., SouthE. Asian Stud. 5: 97. 1967; Tagawa & K.Iwats., Fl. Thailand 3: 368. 1988; Boonkerd & Pollawatn, Pterid. Thailand: 163. 2000; Newman et al., Checkl. Vasc. Pl. Lao PDR: 32. 2007. – Type: Wallich 379, Nepal.

Sagenia membranifolia auct. non Christ: Hosseus, Beih. Bot. Centralbl. 28(2): 366. 1911.

Tectaria brachiata auct. non (Zoll. & Moritzi) Morton: Newman et al., Checkl. Vasc. Pl. Lao PDR: 32. 2007.

Description

Rhizome short, creeping, ascending or erect, 35 mm diam.; scales linear, hairy at margin, rather stiff, bicoloured by nearly black central portion with brown ferrugineous edges or concolorous brown, up to 7 by 0.8 mm. Stipes pale brown to castaneous, 15–35 cm in sterile and up to 60 cm in fertile fronds, densely pubescent on adaxial surface, glabrous beneath. Laminae ovate-subdeltoid or pentagonal, up to 40 by 30 cm, tripinnatifid at base; lateral pinnae 2–4 pairs, the basal pinna much the largest, stalked, asymmetrically subtriangular, acute at apex, with one or two basal basiscopic pinnules; upper pinnae shortly stalked, deeply lobed or with a free sessile basal basiscopic pinnule; apical pinna subdeltoid, cuneate and a little decurrent at base, deeply lobed to pinnatifid; herbaceous, green, glabrous on laminar surface; rachis and pinna-rachis with dense articulated hairs above, glabrous beneath; veins forming copious anastomosis with included veinlets. Sori terminal on free included veinlets, round, usually in a single row at each side of midrib, more or less raised on upper surface; indusia up to 1 mm diam., persistent, glabrous .

Distribution in Thailand

Throughout Thailand, especially on limestone.

Distribution in Laos

Champasak, Khammouane, Xieng Khouang.

Distribution in Cambodia

Kampot, Mondulkiri, Pursat.

Wider Distribution

N India, SW China, Indochina, Taiwan, W Malaysia to Java.

Ecology

On rather dry mountain slopes in not so dense forests at low to medium altitudes up to 900 m alt., especially on limestone, fairly common throughout Thailand.

Proposed IUCN Conservation Assessment

Least Concern (LC). This species is common and widespread and not under any known threat.

Voucher specimens - Thailand

Middleton et al. 4605, Trat, Ko Chang, Thanmagon Waterfall (E).

Voucher specimens - Laos

Maxwell 97-1165, Champasak (CMU); M.F. Newman LAO 191, Khammouane, Nam Theun (E).

Voucher specimens - Cambodia

Long et al. CL407 & CL476, Mondulkiri (P).