Aromatic perennials usually tall and robust,
but sometimes scrambling. Leaves 1–3-pinnate or -ternate, ultimate segments
usually large and distant; petiole greatly to moderately winged at base
. Umbels compound, rays and pedicels 10 to numerous; bracts few or none;
bracteoles several, linear. Calyx teeth minute or obsolete. Petals obovate,
emarginate. Stylopodium flat-domed, flanged at the margin (especially in
young fruit). Fruit broadly ellipsoid to orbicular and notched at both
ends, greatly dorsally compressed with lateral ribs broadly winged, mericarps
with dark vittae lines between the five dorsal ribs.
Key to species
||Rather weak-stemmed almost glabrous plant, usually scrambling;
umbels with slender rays 2–5.5cm long
||Large, sturdy erect plants, at least inflorescence branches
pubescent; umbels with stout rays up to 15cm
||Leaflets with distinct petiolule, glabrous below, petiole
wings of upper leaves not greatly enlarged
||Leaflets usually decurrent into the leaf rachis, densely
puberulent on the veins below, petiole wings of upper leaves greatly enlarged
often 6–11cm broad
sikkimensis (C.B.Clarke) P.K.Mukherjee
Peucedanum sikkimense C.B.Clarke
Slender branched, glabrous diffuse herb, stems weakly erect
or scrambling to 1–2m, plant often tinged purple. Leaves ovate in outline,
distantly 2-pinnate, 18–40 x 8–25cm; ultimate segments ovate-lanceolate
to lanceolate, acuminate, 6–30(–50) x 3–8(–12)mm, petiolulate, base cuneate,
margins coarsely serrate; petiole only narrowly winged at base. Umbels
very diffusely branched with 6–11(–16) spreading rays, to 15cm across;
rays very slender, 2–5.5cm; bracts absent or 1–2, small and like the upper
leaves; umbellules 1–1.5cm across, 7–13-flowered; bracteoles absent or
1–2, linear-lanceolate 2–6mm; pedicels filiform, unequal, 3–8mm, elongating
to 14mm in fruit. Calyx teeth minute triangular, c 0.2mm. Petals white–purplish
or dull yellow, 0.5–1mm. Fruit ellipsoid 6–7.5 x 5–6mm, broadly (1.5–2mm)
winged along both margins, dorsal ridges scarcely prominent, centre dark
brown, wings and ridges paler.
Bhutan: S—Deothang district (Deothang to Wamrong);
C—Ha district (Damthang) and Thimphu district (Chapcha to Confluence,
Dechhenphu, Dolena, Paga, Paro, Thimphu)
Sikkim: Kalej Khola, Lagyap, Lingdam, Rishi, Tsomgo.
Ecology: Coniferous and mixed deciduous forests,
open hillsides, 1200–3300m. Flowers July–August(–September).
Illustration [Fig. 52 f,g]
Specimen List 
Note: Apparently endemic
to Sikkim and Bhutan.
nubigena (C.B.Clarke) P.K.Mukherjee
Heracleum nubigenum C.B.Clarke.
Plants with glabrous stout stems to 1–2m high, thick rhizome
long and tapering. Leaves broadly ovate in outline, 3-pinnate, lower leaves
c 40 x 35cm; leaflets ovate 2.5–7.5 x 1.4–4cm often ternately lobed, margins
serrate, lamina sparsely puberulent above especially on the main veins,
glabrous or almost so below, petiolulate; petioles winged but not greatly
so. Primary umbels large, 17–32cm across, 30–50-rayed, overtopped by lateral
umbels later in the season; peduncles increasing in pubescence towards
the flower head; rays stout, 5–15cm, densely pubescent; umbellules 15–30-flowered;
pedicels 5–15mm, densely pubescent; bracteoles 4–10, linear 5–12mm. Calyx
teeth obsolete. Flowers pale yellow-green. Fruit orbicular 5–8 x 5.5–7mm,
notched at both ends, dorsal ridges broad almost obscuring the dark vittae
Sikkim: Cho La and Yak La. 3000–4200m. Flowers
Specimen List 
Note: The specific
limits between this and the following species is unsatisfactory and need
further study across the whole Sino-Himalayan region.
cyclocarpa (C.Norman) Cannon
Archangelica cyclocarpa C.Norman.
Similar to A. nubigena, but rather larger to 3.5m and with
conspicuous enlarged petiole wings of the upper leaves. Lower leaves to
1m; ultimate segments ovate-lanceolate or ternately-lobed, 4–15 x 2–6cm,
base narrowed and often decurrent on rachis, margin coarsely serrate, lamina
sparsely pubescent above, densely so on the veins below, rachis sparsely
pubescent denser around the nodes; petioles of upper leaves with very wide,
pale wings (3–)6–11cm broad. Fruit 5.5–7 x 5.5–6.5mm, dorsal ridges broad
almost obscuring the dark vittae lines.
Bhutan: C—Thimphu district (Doche La)
Sikkim: Lachen, Tenehungkar, Yumthang
Ecology: Mixed forests and amongst shrubs in open
areas, 2800–4000m. July–September.
Specimen List 
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