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Cortiella C.Norman
Mark F. Watson
Royal Botanic Garden Edinburgh

Acaulous or shortly stemmed perennials, usually forming compact flat rosettes closely adpressed to the soil surface; vertical tap-root long and stout, crowns surrounded by fibrous leaf remains; leaf rachis and main veins, petioles and inflorescence branches stout, densely puberulent to almost glabrous. Leaves ovate-oblong in outline, pinnate; pinnae usually not disjunct on the rachis, 2 x pinnatifid, ultimate segments linear-lanceolate. Umbels compound, main inflorescence usually sessile, laterals often stalked; bracts leaf-like; bracteoles linear-lanceolate or pinnatifid with linear segments. Calyx teeth linear-lanceolate, unequal. Petals narrowly obovate, acuminate at the base, acute apex inflexed. Stylopodium domed, styles long, spreading in fruit. Fruit rectangular-ellipsoid to suborbicular in outline, truncate or more usually cordate at ends, mericarps dorsally compressed, white with 4 dark brown lines of vittae alternating with the 3 dorsal ribs; lateral ribs broadly winged, some of the dorsal ribs equally broadly winged or wings reduced to abortive structures at the base of the fruit; whole fruit characteristically asymmetric with one mericarp having the 2 outer dorsal ribs ornamented and the other only the centre rib, vittae apparent as brown lines.
Key to species
1 Ultimate leaf segments <4mm long; styles long >1.5mm after flowering; wings on dorsal ribs of mericarp well developed, often convoluted and crowded obscuring vittae lines C. hookeri
+ Ultimate leaf segments usually >4mm long; styles shorter; wings on dorsal ribs of mericarp poorly developed, often reduced to the lower half of the fruit or less C. cortioides

Cortiella hookeri (C.B.Clarke) C.Norman
    Cortia hookeri C.B.Clarke p.p.
    Cortia cauwetmarciana M.A.Farille & S.B.Malla
    Cortia glacialis Bonner
Plants 7–30cm across. Leaves 3–10 x 0.8–2cm (including petiole), ultimate segments 2–4 x 0.75mm. Peduncles of lateral umbels 2–10cm, crassicaule 1.5–5mm thick; umbels 7–many-rayed; rays 1–7cm; umbellules 1–1.5cm across, pedicels 2–5mm. Calyx teeth linear-subulate, 0.3–1(–1.2)mm. Petals usually white, sometimes purplish, 1.5–2mm. Styles (1.5–)2–3.5mm after flowering. Fruit 3–5 x 3–5mm, occasionally tipped purple, wings on dorsal ribs of mericarp well developed equalling the laterals (1mm wide), often convoluted and crowded obscuring dark vittae lines at maturity. 

Bhutan: C—Tongsa district (Rinchen Chu, Takse La); N—Upper Mo Chu district (Laya, Yale La), Upper Pho Chu district (Kesha La), Upper Bumthang Chu district (Marlung) and Upper Kuru Chu district (Gong La).
Chumbi: Phari Dzong, Tang La.
Sikkim: Dongkya La, Dzongri, Khangchengyao, Lhonak, Naku La, Momay Samdong. 

Ecology: Alpine turf and particularly screes, 3950–5500m. Flowers June–September. 

Illustration [Fig. 51 i,j]

Specimen List [4101]

Cortiella cortioides (C.Norman) M.F.Watson
    Selinum cortioides C.Norman
    Cortia hookeri C.B.Clarke p.p.
Plants 7–26cm across. Leaves 8–14 x 2–4cm (including petiole), ultimate segments (3–)4–13 x 0.75mm. Main umbel sessile on crown of plant, many-rayed; lateral secondary umbels on peduncles, sometimes crassicaule, to 10cm long, 1.2–2.5mm thick, fewer rayed; rays 0.8–4cm, somewhat shorter than leaves, spreading almost horizontally; umbellules –1.5cm across, pedicels 3–6mmm; bracteoles often tipped purple-black. Calyx teeth 0.3–0.7(–1)mm. Petals white occasionally tinged purple, 1–1.5 x 0.6–0.75mm. Styles 1.75–3mm after flowering. Fruit 4–5.5 x 4–5mm, 3 dorsal ribs prominent, some with reduced wings up the lower half (rarely more) to tiny abortive structures the very lowest part. 

Bhutan: C—Thimphu district (Dungtso La); N—Upper Mo Chu district (Lingshi La), Upper Mangde Chu district (Gyophu La) and Upper Kuru Chu district (Narim Thang).
Chumbi: Kalaeree, Upper Khanghu.
Sikkim: Bikbari, Dzongri, Ghora La, Goecha La, Jelep La, Khangchengyao Glacier, Kongra La, Lhonak, Phalut, Riago, Samiti Lake, Yale La, Yume Samdong. 

Ecology: Screes, rocky slopes and free draining sandy areas, 4000–5400m. Flowers (June–)July–September(–October). 

Specimen List [15151]

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